Abstract
Targeted gene silencing by RNAi requires the RNA-induced silencing complex (RISC), whose core component is the protein Argonaute (Ago) bound to a microRNA (miRNA) or an siRNA. In humans, Ago2 is loaded with miRNAs by the action of a specialized assembly called the RISC-loading complex (RLC), comprising the proteins Ago2, Dicer, and TRBP. Here we show that the human RLC assembles spontaneously in vitro from purified components. No cofactors or chaperones are required for the complex to form. The reconstituted RLC, containing one copy of each protein, has the dicing, slicing, guide-strand selection, and Ago2-loading activities observed for the endogenous RLC. Furthermore, once Ago2 is loaded with an miRNA, it tends to dissociate from the rest of the complex. These results lay the groundwork for future structural and functional dissection of RISC loading in humans.
MeSH Terms
Animals
Argonaute Proteins
Carboxypeptidases/chemistry
Catalysis
Cell Line
DEAD-box RNA Helicases/metabolism
Drosophila
Endoribonucleases/metabolism
Eukaryotic Initiation Factor-2/metabolism
Gene Silencing
Humans
Mass Spectrometry/methods
MicroRNAs/metabolism
Models, Biological
Protein Conformation
RNA/chemistry
RNA Interference
Ribonuclease III
Chemicals
AGO2 protein, human
Argonaute Proteins
Eukaryotic Initiation Factor-2
MicroRNAs
RNA
Endoribonucleases
DICER1 protein, human
Ribonuclease III
Carboxypeptidases
SCPEP1 protein, human
DEAD-box RNA Helicases
Authors & Affiliations
5 authors, click to expand affiliations / ORCID
MacRae Ian J
Department of Molecular and Cell Biology, University of California, Berkeley, CA 94720, USA. macrae@scripps.edu
Ma Enbo
Zhou Min
Robinson Carol V
Doudna Jennifer A
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